European paper wasps are black wasps with yellow markings including rings on their body. They also have spots on their face and body which show information about gender. The spots might also have information about social status and whether they are aggressive. Faces with more black might show dominance and give information about the size of the wasp. European paper wasps have bright reddish-orange antennae. (Buck, et al., 2008a; Buck, et al., 2008b; Buck, et al., February 19, 2008; Green and Field, 2010)
European paper wasps look like Northern paper wasps, but they are smaller, about 2.0 cm long on average. They have 6 legs, a pair of antennae, and a pair of wings. They have an indentation on the side of the middle part of their bodies, and ridges on the front of their abdomens. They are different from closely related species because they fly with their back legs slightly below their bodies. Females are larger than males, and their undersides are black instead of yellow. Front wings are 9.5 to 13.0 mm long in females, and 8.5 to 12.0 mm long in males. (Buck, et al., 2008a; Buck, et al., February 19, 2008; Cranshaw, 2008)
European paper wasps originally lived in the Mediterranean, northern Africa, southern Europe, the Middle East, and eastern China. They were introduced to North America in the 1970s and 1980s. At first, they were mostly found on the East coast of the United States. They have spread into the Midwest and the western and southwestern United States. European paper wasps were introduced to Canada in the 1990s, including Ontario, Nova Scotia, and British Columbia. They have also been introduced to Chile and Argentina and also western Australia. (Cervo, et al., 2000; Jacobs, 2011)
European paper wasps live on land in forests, grasslands, and in the hot and dry chaparral. They live in temperate climates, meaning places where the seasons change. They are found in urban, suburban, and agricultural areas. They tend to be located close to human civilization because they nest in things humans build. They also live in forests and on plants where they can feed and nest. When they are nesting, they choose spaces made by farm machines or things used for recreation. In the winter, pregnant queens stay in protected areas like in the walls of houses or hollow trees. Then, females build nests in these places or somewhere nearby at the beginning of spring. (Buck, et al., February 19, 2008; Jacobs, 2011; Rusina, et al., 2011)
European paper wasp eggs grow into larvae. Queen and worker wasps collect food for the larvae. The larvae go through a pupa state, and then transform into adults. At the right temperature and with enough food, adults take 40 days to develop into adults. Adults drum with their antennae on the larvae, which determines whether the larvae grow up into workers or founder wasps. (Suryanarayanan, et al., 2011)
European paper wasp colonies have a queen who is head of the colony. The queen mates with more than one male, and is more likely to mate with resident males than transient males. Resident males defend a large tree or other spot for multiple days, while transient males move between defending different places. To choose mates, the queen flies over the territories of the males. She probably chooses a resident male because they are often large and aggressive. After the larvae develop, workers help the foundress female provide food to the other larvae. (Beani and Turillazzi, 1987; Beani, et al., 1992)
Queen European paper wasps are more likely to mate with males that don't share their nest. Males don't treat females differently if they are from the same nest. Females are picky in mating. (Liebert, et al., 2010)
Female European paper wasps mate during the spring and have more than one set offspring. They mate once more in the fall, and store male reproductive cells in their bodies until spring. They spend the winter in houses, walls, or outdoors. In April and May, they start to build nests in places that are dark and protected. They nest in places built by humans more often than northern paper wasps. Females nest alone or in combined nests with other females. Once in a while, females stay in the place where they hibernate until they take over an abandoned nest. Nests are built from papery materials and attached to wood, metal, or rock. At the beginning of the season, females lay eggs in the paper holes of the nest. During the season, worker wasps rebuild the nest and make it bigger. (Cranshaw, 2008; Zanette and Field, 2011)
Females lay eggs in the holes of the nest, which grow into adults in 3 to 4 weeks. The eggs hatch into larvae in 3 to 5 days. Then, the queen wasp feeds them chewed-up caterpillars. The larvae close off the top of a nest cell with silk and change from pupae to adults. As adult wasps develop, they become worker wasps in the colony. By the summer, the colony has several dozen wasps. They help the foundress get food for future larvae and build space in the nest for them. Later on in the season, some of the workers reproduce. Only females survive the winter. (Cranshaw, 2008)
Female European paper wasps protect and feed their offspring until after they are able to feed themselves. The eggs develop inside their bodies with yolks, and females keep the eggs in her body until she lays them in a nest. Then, the foundress of the colony feeds her larvae chewed up insects like caterpillars. The first adults become worker wasps who help build the nests and collect food for the next group of young. (Bartelt, 2011)
Queen European paper wasps live longer than workers because queens are protected by females and hibernate during the winter. Colonies start in late March or April, and can have more than one queen. Only females that reproduce survive the winter, so most European paper wasps live less than one year. Females that reproduce can survive more than one season. (Cranshaw, 2008; Strassmann, et al., 2004)
European paper wasps are able to fly and walk. They are social wasps that live in large groups called colonies. Each colony has a hierarchy between the queen and workers. There are also separate hierarchies between the working wasps. Workers spend their time moving around the nest, hunting, building up the nest, and sitting on the nest. European paper wasps are active in the day and come back to the nest at night. Males are sometimes aggressive, and their level of aggression is related to the pattern on their face. Males might be more likely to challenge a weaker rival based on the pattern on its face. (Baracchi, et al., 2010; Cranshaw, 2008)
Female queens hibernate during the winter and lay eggs in early spring. Eggs become workers for the colony and the queen. Together, they build and live in a nest. The female foundresses of the colonies visit fewer areas of the nest than the workers do. By the summer, the colony has several dozen wasps. The wasps born first often have higher social status, and all the workers add to the nest as the colony gets bigger. In smaller colonies, the 6 wasps born first have less interaction with highly ranked wasps. In the next season, females usually choose a spot to build a new nest. (Baracchi, et al., 2010; Cranshaw, 2008; Karsai, et al., 1996)
Scientists don't know much about the size of the home range of European paper wasps.
European paper wasps might not be able to recognize each other as specific individuals. Scientists know this because they are just as aggressive with each other if they have met each other before or not. To figure out if another wasp belongs to their nest or not, they use chemicals on the outside waxy layer of their bodies. (Bruschini, et al., 2011; Sheehan and Tibbetts, 2010)
European paper wasps use touch to communicate dominance. In a behavior called mounting, the dominant wasp drums their antennae on on another wasp's head, and the other wasp lowers its head and antennae. In a behavior called boxing, the wasp uses its front legs to bat at another wasp. They curl up together and the wasp trying to be dominant wraps its body around the other one. European paper wasps also lightly bat the heads of other wasps with their antennae or drum on their bodies apart from struggling for dominance. (Izzo, 2011)
European paper wasps are omnivores, so they eat animals and plants. They eat insect larvae and caterpillars, as well as aphids, honeydew melon, and nectar from flowers. European paper wasps consume more kinds of insects than northern paper wasps, which only eat caterpillars. (Cervo, et al., 2008)
European paper wasps look similar to closely related species, so potential predators may have trouble telling different kinds of wasps apart. (Bartelt, 2011)
European paper wasps help pollinate plants. They also eat insects, especially caterpillars. Where European paper wasps have been introduced, they affect native northern paper wasps. European paper wasps set up their nests before native wasps, so they can increase the size of their colony without a lot of competition. Nesting early also helps them avoid bird predators. European paper wasps eat a bigger variety of insects than native species, which is good for the health of the water. Their success makes it more difficult for native wasps to survive. (Cervo, et al., 2000; Cranshaw, 2008; Jacobs, 2011; Stahlhut, et al., 2006)
Native European paper wasps get infected by parasitic insects that are called parasitoids, such as Endurus argiolus and Xenos vesparum. Adults lay eggs on or inside European paper wasps. They larvae that infect sometimes kill their hosts. In North America, native parasitoids don't recognize European paper wasps as hosts. In Italy and the northeastern United States, European paper wasps can be infected with a bacteria parasite called These North American parasitoids do not generally infect European paper wasps in the United States because they do not recognize them as hosts based on their recent introduction. In Italy and the northeastern United States, European paper wasps can get infected with Wolbachia bacteria. (Cervo, et al., 2000; Jacobs, 2011; Stahlhut, et al., 2006)
European paper wasps sting people and other animals who come too close to their nests. They can also be threatened by native wasps. European paper wasps nest in structures or buildings built by humans more often than their closest relatives. This means people kill them with chemicals and get rid of their nests. (Bartelt, 2011; Buck, et al., 2008a)
European paper wasps benefit humans by pollinating plants. They help reduce the number of insects like caterpillars, hornworm larvae, cabbageworms, tent caterpillars, and sawfly larvae like Cimbicidae, Diprionidae, and Tenthredinidae. (Bartelt, 2011; Cranshaw, 2008)
European paper wasps are not threatened or endangered.
Eliza Stout (author), University of Michigan-Ann Arbor, Catherine Kent (editor), Special Projects.
Living in Australia, New Zealand, Tasmania, New Guinea and associated islands.
living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.
living in the southern part of the New World. In other words, Central and South America.
living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.
living in landscapes dominated by human agriculture.
having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends.
an animal that mainly eats meat
mid-altitude coastal areas with mild, rainy winters and long, dry summers. Dominant plant types are dense, evergreen shrubs.
uses smells or other chemicals to communicate
used loosely to describe any group of organisms living together or in close proximity to each other - for example nesting shorebirds that live in large colonies. More specifically refers to a group of organisms in which members act as specialized subunits (a continuous, modular society) - as in clonal organisms.
helpers provide assistance in raising young that are not their own
ranking system or pecking order among members of a long-term social group, where dominance status affects access to resources or mates
animals which must use heat acquired from the environment and behavioral adaptations to regulate body temperature
the condition in which individuals in a group display each of the following three traits: cooperative care of young; some individuals in the group give up reproduction and specialize in care of young; overlap of at least two generations of life stages capable of contributing to colony labor
parental care is carried out by females
union of egg and spermatozoan
forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.
an animal that mainly eats fruit
An animal that eats mainly plants or parts of plants.
the state that some animals enter during winter in which bodily functions slow down, reducing their energy requirements so that they can live through a season with little food.
An animal that eats mainly insects or spiders.
fertilization takes place within the female's body
referring to animal species that have been transported to and established populations in regions outside of their natural range, usually through human action.
offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).
A large change in the shape or structure of an animal that happens as the animal grows. In insects, "incomplete metamorphosis" is when young animals are similar to adults and change gradually into the adult form, and "complete metamorphosis" is when there is a profound change between larval and adult forms. Butterflies have complete metamorphosis, grasshoppers have incomplete metamorphosis.
having the capacity to move from one place to another.
the area in which the animal is naturally found, the region in which it is endemic.
an animal that mainly eats nectar from flowers
an animal that mainly eats all kinds of things, including plants and animals
reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.
Referring to a mating system in which a female mates with several males during one breeding season (compare polygynous).
breeding is confined to a particular season
reproduction that includes combining the genetic contribution of two individuals, a male and a female
associates with others of its species; forms social groups.
living in residential areas on the outskirts of large cities or towns.
uses touch to communicate
that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).
Living on the ground.
A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.
living in cities and large towns, landscapes dominated by human structures and activity.
movements of a hard surface that are produced by animals as signals to others
uses sight to communicate
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